"My father left his assets in two trusts -- for the benefit of myself and my sibling. His best friend is the trustee who has delegated all responsibilities to both an investment house and high-end law firm. He takes a .4% fee -- with all fees now totaling 2.25% to 3%. What is legal? What is moral?"
As for the moral issue, that's tough to call without understanding what the trustee does. You've also mentioned nothing about the assets in the trusts, the rates of return realized, whether extra services were performed in a particular year, and so on. However, the moral obligation of the trustee should be to carry out your father's wishes. That should be the guiding principal for everyone involved. On the flip side would your father want you to sue a lifetime close friend?
Perhaps the best approach is to request a meeting with the trustee and present your concerns. Considering that investment advisers and financial planners often suggest a withdrawal rate from a trust in the order of 3-5% and many states Principal and Income Acts (PIA) provide for elections to make distributions based on those percentages of trust assets, the overall fee structure could have a negative impact on the economics of the entire trust. So you're right to address the issue.
Before meeting with the trustee meet with a trust attorney in the state whose laws govern the trust and review the trust document, fee structure and applicable state law. You want to be prepared before the meeting (even if you meet informally without lawyers).
If the trustee won't reasonably address your concerns, then you'll have to hire an attorney to address the issues with the trustee. But if the trustee is being advised by a "high-end" law firm presumably he or she has documented the basis for the fees involved.